|Ordered||10 Apr 1941|
|Laid down||28 Oct 1942||Deutsche Werke AG, Kiel (werk 309)|
|Launched||17 Jul 1943|
|Commissioned||8 Sep 1943||Oblt. Rudolf Rademacher|
|Successes||No ships sunk or damaged|
Sunk 30 June, 1944 north-east of the Faeroes, in position 63.27N, 00.50W, by depth charges from Canadian Canso (RCAF-Sqdn. 162/A) and British Liberator aircraft (Sqdn. 86/E). 52 dead (all hands lost).
Men lost from U-boats
Unlike many other U-boats, which during their service lost men due to accidents and various other causes, U-478 did not suffer any casualties (we know of) until the time of her loss.